Listing of names which have been applied in the genus Kiefferulus Goetghebuer
(includes the previous genera Nilodorum Kieffer and Carteronica Kieffer)

Compiled by Jon Martin, Biosciences 5, School of Biosciences, University of Melbourne, Victoria, 3010, Australia.

This listing recognizes Kiefferulus in the wider sense as established by Cranston, Webb, & Martin (1989)

format: LINE a. NAMES (HOMONYMS preceded by a number). $ = VALID NAME or SYNONYM * = NOW IN ANOTHER GENUS. # = STATUS UNCERTAIN. LINE b. PRESENT TAXONOMIC STATUS WITH SOURCE OF INFORMATION (Not necessarily authority for designation). LINE c. ORIGINAL DESCRIBER & DATE (Original genus also, if not KIEFFERULUS). LINE d. DISTRIBUTION & TYPE LOCALITY. LINE e. OTHER REFERENCES.

ABCDEFGHIJKLMNOPQRSTUVWXYZ

A

a. $ ALBITARSE
b. NEW NAME: RUFOSUM - FREEMAN 1957A: 378 - as C. (NILODORUM).  CRANSTON, WEBB & MARTIN 1989: 420 (generic position).
c. KIEFFER 1922I: 46 - as NILODORUM. Transfered to CHIRONOMUS (NILODORUM) by FREEMAN 1957A: 378, so name preoccupied by LUNDSTROEM 1913.
AFRICA - Shambe, SUDAN (Type lost?).
FREEMAN 1956B : 94 - as NILODORUM.  FREEMAN & CRANSTON 1980A: 194 - as NILODORUM RUFOSUM.

a. $ ATROXITARSIS
b. KIEFFERULUS
c. DAS & MAZUMDAR 2012: 1 - as KIEFFERULUS, on basis of all life stages.
d. INDIA - Mirik (26.53, 88.12), West Bengal.
e.

B

a. * BRASILIENSIS
b. Nomen dubium in CHIRONOMINAE - SPIES & REISS 1996: 74.
c. WIEDEMANN 1828A: 15 - as CHIRONOMUS, on basis of adult male?
d. SOUTH AMERICA - Montevideo, URUGUAY; ARGENTINA.
e. LYNCH ARRIBALZAGA 1893A: 253 - Redescription.   KIEFFER 1906C: 16 - as CHIRONOMUS. EDWARDS 1931A: 320 - as CHIRONOMUS (NILODORUM) BRAZILIENSIS.   OLIVEIRA 1954A: 453 - Redescription. CRANSTON, WEBB & MARTIN 1989: 420 - not KIEFFERULUS, but uncertain whether true NILODORUM.

a. * BRAZILIENSIS
b. Invalid name.
c. EDWARDS 1931A: 320 - as CHIRONOMUS (NILODORUM), misspelling of BRASILIENSIS WIEDEMANN.
d. SOUTH AMERICA - URUGUAY.
e. SPIES & REISS 1996: 74 - as incorrect spelling of BRASILIENSIS.

a. $ BREVIBUCCA
b. KIEFFERULUS - CRANSTON, WEBB & MARTIN 1990: 421.
c. KIEFFER 1922: 45 - as NILODORUM, on basis of adult. Transferred to CHIRONOMUS (NILODORUM) by GOETGHEBUER 1936F?d. AFROTROPICAL - Shambe, SUDAN; ASIA, ISRAEL, EGYPT.
e. GOETGHEBUER 1937-54 (13c): 34 - as TENDIPES (NILODORUM). FREEMAN 1955B: 19 - as NILODORUM. FREEMAN 1957A: 375 - as CHIRONOMUS (NILODORUM). ASHE & CRANSTON 1990: 291 - as NILODORUM.

a. * BREVIPALPIS (BREVIPALPE)
b. KIEFFERULUS - CRANSTON, WEBB & MARTIN 1990: 421.
c. KIEFFER 1918B: 69 - as CHIRONOMUS, on basis of adult male and female.
d. AFRICA - Lac Dembel, ETHIOPIA.
e. FREEMAN 1957A: 376 - as CHIRONOMUS (NILODORUM). FREEMAN & CRANSTON 1980A: 194 - as NILODORUM.

a. * BREVIVALVA
b. SYN: Nomen dubium in CHIRONOMINAE - SPIES & REISS 1996: 74.
c. KIEFFER 1917B: 351 - as CHIRONOMUS, on basis of adult male.
d. SOUTH AMERICA - PARAGUAY.
e. EDWARDS 1931A: 300 - as syn. CHIRONOMUS (NILODORUM) BRAZILIENSIS. OLIVIERA 1954A: 453 - SYNONYMY.

a. $ BURGEONI
b. SYN: BREVIBUCCA - FREEMAN 1957A: 375 (as C. (NILODORUM)). CRANSTON, WEBB & MARTIN 1990: 421 (Gen.posn.).
c. GOETGHEBUER 1935E: 398 - as NILODORUM. Transferred to CHIRONOMUS (NILODORUM) by GOETGHEBUER 1936F: 464.
d. AFRICA - Kamande,ZAIRE.
e.FREEMAN & CRANSTON 1980A: 193 - as syn NILODORUM BREVIBUCCA.

C

a. $ COCLEARIS
b. KIEFFERULUS
c. DAS & MAZUMDAR 2012: 7 - as KIEFFERULUS, on basis of all life stages.
d. INDIA - Mirik (26.53, 88.12), West Bengal.
e.

a. $ 'CORNISHI'
b. KIEFFERULUS
c. MARTIN - manuscript name, as KIEFFERULUS, on basis of cytology.
d. AUSTRALIA - Werribee, Victoria.
e. MARTIN, HOFFMAN & CASE 1996: 470 - photograph of chromosomes.  MARTIN, BLINOV, ALIEVA & HIRABAYASHI 2007:195 - molecular phylogeny.

D

a. $ DISPARILIS
b. KIEFFERULUS - CRANSTON & JUDD 1989: 255.
c. GOETGHEBUER 1936F: 462 - as ?GLYPTOTENDIPES. Transferred to CHIRONOMUS (ENDOCHIRONOMUS ) by FREEMAN 1957A: 353.
d. AFRICA - Parc National Albert, ZAIRE.
e. FREEMAN 1955B: 24; FREEMAN & CRANSTON 1980A: 191 - as ENDOCHIRONOMUS. HARRISON 1987: 41 - as EINFELDIA. CRANSTON, WEBB & MARTIN 1990: 421 - as KIEFFERULUS.

a. $ DUX
b. KIEFFERULUS - BECK & BECK 1970A: 38.
c. JOHANNSEN 1905A: 231 - as CHIRONOMUS.
d. NEARCTIC - Ithaca, New York, U.S.A.
e. JOHANNSEN 1937B: 45 - as CHIRONOMUS (CRYPTOCHIRONOMUS). TOWNES 1945A: 110 - as TENDIPES (KIEFFERULUS) . SUBLETTE & SUBLETTE 1965A: 170 - as CHIRONOMUS (KIEFFERULUS) . CRANSTON, WEBB & MARTIN 1990: 421 - as KIEFFERULUS.

E

F

G

H

I

a. $ INCIDERUS
b. KIEFFERULUS
c. CHATTOPADHYAY & CHAUDHURI 1991: 24 - as KIEFFERULUS, on basis of adult male.
d. ASIA - Bankura, West Bengal, INDIA.
e.

L

a. * LATILOBUS
b. SYN: KIEFFERULUS CHLORONOTUS - FREEMAN 1957A: 371 (as CH. (KIEFFERULUS)).  FREEMAN & CRANSTON 1980A: 191 (generic position).
c. KIEFFER 1923A: 384.
d. AFRICA - Salisbury, S.RHODESIA (ZIMBABWE).
e. FREEMAN 1955D: 371 - as CHIRONOMUS.

a. $ LATIVALVA (LATIVALVUS)
b. SYN: KIEFFERULUS INTERTINCTUS - FREEMAN 1961A: 689 (as CHIRONOMUS (KIEFFERULUS)). MARTIN 1973A: 1372 (Generic position).
c. KIEFFER 1917A: 211.
d. AUSTRALIA - Sydney, NEW SOUTH WALES.
e. EDWARDS 1929A: 375 - as PENTAPEDILUM (KIEFFERULUS).  CRANSTON, WEBB & MARTIN 1990: 421 - as syn. K. INTERTINCTUS.

a. $ LONGILOBUS
b. KIEFFERULUS - CRANSTON, WEBB & MARTIN 1990: 421.
c. KIEFFER 1916B: 107 - as TENDIPES, on basis of adult male and female.
d. OCEANIA - Takao, FORMOSA; MICRONESIA; AUSTRALIA; OKINAWA; Thailand.
e. KIEFFER 1921M: 591 - as CARTERIA.  STRAND 1928A: 48 - as CARTERONICA.  TOKUNAGA 1940A: 220; 1964A: 552 (Redescription, but was larva correctly associated?); SUBLETTE & SUBLETTE 1973B: 402; SASA & HASEGAWA 1983: 313 - as CHIRONOMUS.  FREEMAN 1957A: 356 - as DICROTENDIPES.  ASHE 1983: 15 - generic position.  CRANSTON 2007: 240 - larval head characters.

M

a. $ MARTINI
b. KIEFFERULUSc. FREEMAN 1961A: 690 - as CHIRONOMUS (DICROTENDIPES), on basis of adult female.
d. AUSTRALIA - Echuca, Vic.
e. MARTIN 1963A: cytology & larval morphology; 1964A: 285 - key.  WEBB & MARTIN 1987: 419 - ultrastructure of ventromentum.  MARTIN, BLINOV, ALIEVA and HIRABAYASHI 2007 - molecular data.

N

a. $ NODULOSUS
b. KIEFFERULUS - CRANSTON, PERS. COMM.
c. HASHIMOTO (in HASHIMOTO, WONGSIRI, WONGSIRI, TIRAWAT, LEWVANICH & YASUMATSU) 1981: 10 - as CHIRONOMUS, EINFELDIA group, on basis of adult male.
d. ASIA - Ban Bangkanark, Chachoengsao Province, THAILAND.
e.

O

a. $ OBSCURATUS
b. SYN : KIEFFERULUS DUX - TOWNES 1945A: 110 (as TENDIPES (KIEFFERULUS)). BECK & BECK 1970A: 38 (generic position).
c. MALLOCH 1915A: 479 - as .
d. NEARCTIC - Dubois, Illinois, U.S.A.
e. FRISON 1927A: 168 - designation of Lectotype. SUBLETTE & SUBLETTE 1965A: 170 - as syn. CHIRONOMUS (KIEFFERULUS) DUX.  WEBB 1980A: 85.

a. $ OPALENSIS
b. KIEFFERULUS
c. FORSYTH 1975A: 215 - as KIEFFERULUS.
d. PACIFIC REGION - Opal Lake, South Auckland, North Island (-38.31, 176.38), NEW ZEALAND.
e. MARTIN, HOFFMAN & CASE 1996: 470 - chromosome photograph.

a. $ OTSURU
b. SYN.: KIEFFERULUS CALLIGASTER - CHAUDHURI & GUHA 1987: 28.
c. SASA & HASAGAWA 1983: 314 - as CHIRONOMUS sp. OTSURU.
d. ASIA - Rykyu Islands, JAPAN.
e.

P

a. $ PAPILLUS
b. KIEFFERULUS
c. DAS & MAZUMDAR 2012: 12 - as KIEFFERULUS, on basis of all life stages.
d. INDIA - Mirik (26.53, 88.12), West Bengal.
e.

a. $ PRUINOSUS (PRUINOSA)
b. KIEFFERULUS - EKREM & STUR 2016.
c. FREEMAN 1961B: 246 - as CHIRONOMUS (ENDOCHIRONOMUS), on basis of adult male.
d. AFRICA - Montagne d'Ambre 1000m., MADAGASCAR NORD.
e. FREEMAN & CRANSTON 1980A: 191 - as ENDOCHIRONOMUS

a. $ PSILOCHIRUS
b. KIEFFERULUS - CHAUDHURI & GUHA 1987: 28.
c. KIEFFER 1911F: 159 - as CHIRONOMUS, on basis of adult male.
d. ASIA - Port Canning, West Bengal, INDIA.
e. SUBLETTE & SUBLETTE 1973B: 414 - as uncertain Chironomini. CRANSTON, WEBB & MARTIN 1990: 421 - as KIEFFERULUS.

a. $ PUNGENS
b. KIEFFERULUS - EPLER 1995: .
c. TOWNES 1945A: 119 - as TENDIPES, on basis of adult male.
d. NORTH AMERICA - Canoe Ck., 12 ml s. St. Cloud, Florida (type), & Little Falls, D.C.(paratype), U.S.A. (April).
e. JOHANNSEN & TOWNES 1952A: 79 - as TENDIPES. EPLER 2001: 8.87 - as K. (WIRTHIELLA).

R

a. $ RENICORNIS
b. KIEFFERULUS
c. DAS & MAZUMDAR 2012: 17 - as KIEFFERULUS, on basis of all life stages.
d. INDIA - Mirik (26.53, 88.12), West Bengal.
e.

a. $ ROSTRATUS
b. KIEFFERULUS - CHAUDHURI & GUHA 1987: 28.
c. KIEFFER 1911F: 164 - as CHIRONOMUS, on basis of adult male.
d. ASIA - Sylhet, ASSAM (Kamrup), INDIA.
e. SUBLETTE & SUBLETTE 1973B: 414 - as unplaced CHIRONOMINI. REE & KIM 1981: 143 - as CRYPTOCHIRONOMUS. CRANSTON, WEBB & MARTIN 1990: 421 - as KIEFFERULUS.

a. $ RUGOSUM
b. KIEFFERULUS - CRANSTON, WEBB & MARTIN 1990: 421.
c. FREEMAN 1957A: 378 - as CHIRONOMUS (NILODORUM) and new name for ALBITARSE KIEFFER.
d. AFRICA - Shambe, SUDAN.
e. FREEMAN & CRANSTON 1980A: 194 - as NILODORUM.

S

a. $ STUPIDUS
b. SYN: BARBATITARSIS - CRANSTON (2002)
c. JOHANNSEN 1932B: 528 - as CHIRONOMUS, on basis of reared adult male & female.  Notes similarity to NILODORUM KIEFFER.
d. INDONESIA - Botanical Garden, Buitenzorg, JAVA.
e. LENZ 1937B (Type of species); SUBLETTE & SUBLETTE 1973B: 409 - asSTICTOTENDIPES.  ALFRED & GEORGE 1990: 183 - Cytology as NILODORUM.

a. $ STYLATUM
b. SYN: BREVIBUCCA - FREEMAN
c. KIEFFER 1922: 46 - as NILODORUM and variety of BREVIBUCCA
d. AFRICA - Shambe, SUDAN
e. ASHE & CRANSTON 1990: 291 - as syn NILODORUM BREVIBUCCA.  CRANSTON, WEBB & MARTIN 1990: 421 - as syn KIEFFERULUS BREVIBUCCA.

a. $ SURDELLUS
b. SYN: KIEFFERULUS BREVIPALPIS - FREEMAN 1957A: 376 (as C.(NILODORUM)).  CRANSTON, WEBB & MARTIN 1990: 421 (generic position).
c. GOETGHEBUER 1936F: 464 - as CHIRONOMUS, on basis of adult.
d. AFRICA - Vitshumbi, ZAIRE.
e. FREEMAN & CRANSTON 1980A: 194 - as syn. NILODORUM BREVIPALPE.

T

a. $ TAINANUS
b. KIEFFERULUS - CRANSTON, WEBB & MARTIN 1990: 421.
c. KIEFFER 1912: 36 - as TENDIPES, on basis of adult male and female.
d. ASIA - Tainan, TAIWAN; JAPAN.
e. KIEFFER 1921M: 586 - as PHYTOCHIRONOMUS.  GOETGHEBUER 1937-54(13c): 13 - asGLYPTOTENDIPES.  LICHTENBERG 1974A: 122 (List of types).  HASHIMOTO 1977B: 84 - asCHIRONOMUS (Unknown subgenus).  SASA 1979B: 6 - as between CHIRONOMUS & EINFELDIA.  SASA& YAMAMOTO 1977A: 312; HASHIMOTO, WONGSIRI, WONGSIRI, TIRAWAT, LEWVANICH & YASUMATSU 1981: 9 - as CHIRONOMUS.  CHAUDHURI & GUHA 1987: 28 - as syn. K. BARBATITARSIS.  CRANSTON & MARTIN 1989: 271; ASHE & CRANSTON 1990: 291 - as NILODORUM.

a. * TENDIPEDIFORMIS (TENDIPEDIFORME)
b. KIEFFERULUS - GOETGHEBUER 1922A: 40 (as type species); FITTKAU, SCHLEE & REISS 1967A: 372.
c. GOETGHEBUER 1921A: 106 - as TANYTARSUS.  Placed in TENDIPES (KIEFFERULUS) by TOWNES 1945A: 110 & in C. (KIEFFERULUS) by FREEMAN 1961A: 689.
d. EUROPE - Destelbergen, Heusden & Gand, BELGIUM; NORTH AFRICA.
e. GOETGHEBUER 1928A: ; EDWARDS 1929A: 375 - as PENTAPEDILUM (KIEFFERULUS).  PAGAST 1936A.  CHERNOVSKII 1949B:. GOETGHEBUER 1937-54(13c): 76 - as section TANYTARSINI, genus KIEFFERULUS.   CRANSTON, WEBB & MARTIN 1990; 421; ASHE & CRANSTON 1990: 286 - as KIEFFERULUS.

a. $ TRIGONUM
b. SYN: KIEFFERULUS NODULOSUS - CRANSTON (PERS. COMM.)
c. SONG, WANG, BU & QI 2020: 72 - as KIEFFERULUS, on basis of adult male.
d. ASIA - Fujian Province, Guizhou Province, Guangxi Zhuang Autonomous Region, and Hainan Province, CHINA.
e.

a. * TUMIDUS
b. SYN: KIEFFERULUS CALLIGASTER - CRANSTON 2002.
c. JOHANNSEN 1932B: 530 - as CHIRONOMUS (PHYTOCHIRONOMUS).
d. S.E.ASIA - JAVA, SUMATRA, INDONESIA.
e. LENZ 1937B: 5 - as TENDOCHIRONOMUS (N.Gen).  SUBLETTE & SUBLETTE 1973B: 404 - as GLYPTOTENDIPES.

U

V

a. $ VITSHUMBIENSIS
b. SYN: KIEFFERULUS BREVIPALPIS - FREEMAN 1957A: 376. CRANSTON, WEBB & MARTIN 1990: 421 (generic position).
c. GOETGHEBUER 1936F: 479 - as CHIRONOMUS.
d. AFRICA - P.N.A.Vitshumbi, ZAIRE.
e. FREEMAN & CRANSTON 1980A: 194 - as syn. NILODORUM BREVIPALPE.