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C. ramosus Chaudhuri et al. 1992 immatures

Larva:  a small to medium plumosus-type (length 8-13.2 mm).   Anterior pair of ventral tubules shorter than posterior pair.  Anal tubules 310 (290-330) µm long.  Gula and frontoclypeus apparently not darkened.
Mentum with c2 teeth well separated and c1 broad (type IB), 4th lateral reduced (type II).
Ventromentum about 3.9 times longer than its depth; figure indicates numerous striae.  Pecten epipharyngis with 20 teeth (type B) - other populations have 12-16 teeth.  Premandible shown with two equal teeth, but normally the inner tooth is 2-3 times wider than the outer tooth.
Antenna with basal segment about 2.25 times longer than wide, RO less than 1/3 up from the base; AR 1.69 (1.53-2.10); ratio of antennal segments 23.1 : 6.7 : 2.2 : 3.1 : 1.6.
Mandible figured as type IIB; no information on number of furrows on outer surface; or number of taeniae in PMa; MTR perhaps about 0.33.


Illustration of larval parts of C. ramosus from Chaudhuri et al.1992

Pupa: Exuvia grey.  Length of male 6.69 (6.67-7.64) mm, of female 7.27 (6.52-8.86).  Frontal tubercles about 0.09 mm long and 0.06 mm wide, with the subapical seta about 0.05 mm in length.  Antennal sheath of female about 540 (540-560) µm long.  Wing sheath 1.54 (1.42-1.72) mm in length, respiratory base elliptical and about 0.15 mm wide.  PSA caudolateral on segment IV-VI; PSB basolateral on segment I and caudolateral on segment II.  Hook row on segment II comprised of about 46-58 hooklets.  Shagreen pattern as shown in figure.  Caudolateral spurs of segment VIII with 3 spines.

Cytology: Four polytene chromosomes with the pseudothummi-cytocomplex combination BF, CD, AE, G.
Nucleolus in arm B (at 11B)(as in map of Nath and Godbole 1997).  Two BRs, about equally distant from each end, are on arm G.  No reported polymorphism.
Some points can be made by reference to the map of Nath & Godbole (1997):
ramA1:    1 - 2c, 10 - 12, 3- 2d, 9 - 4, 13 - 19                as circumdatus A2, holomelas, etc.
ramB1:    Characteristic bands at 10F.  Nucleolus at 11B
ramC1:    Developmentally stage specific puff at 6B.            as C2 of incertipenis?
ramD1:   
ramE1:    1a-i, 5 - 10b, 3e - 2, 4 - 3f, 10c - 13            i.e Inv2-5 from aprilinus, as incertipenis
ramF1:    1 - 2a, 10 - 3d, 14c - 11, 2b - 3c, 14d - 23.
ram G1:    BRs at 18C and 19C

Chromosomes also described by De & Gupta (1994), as C. niger.

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Modified: 17 April 2022
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