Gular region pale in some Japanese populations, slightly dark to dark over posterior 1/3 to 1/2, Frontoclypeus darkened.
Mentum (Fig. c) with c1 tooth tall, c2 teeth well separated (type III), 4th laterals slightly reduced (type I-II).
Ventromentum (Fig. d) with smooth anterior margin and about 38 (34-44) striae; IPD 53-71 µm (0.31-0.40 of mentum width); VMR about 0.24-0.32.
Pecten epipharyngis (Fig. a) with about 12-16 broad sharp teeth (type B).
Premandible with outer tooth slightly longer and coming to a relatively narrow point, inner tooth coming to a broad point and 2-3.7x wider.
Antenna (Fig. b) with basal segment about 3.4 (2.8-4.0) times longer than wide, ring organ about one third to 2/5th up from base; AR about 2.08 (1.94-2.44), segment lengths (microns) 108 : 28.5 : 8.5 : 11 : 6.
Distance between the antennal bases generally greater than that between the S4 setae.
Mandible (Fig. e) with 3rd inner tooth partly or sometimes almost fully separated,and partly darkened (type II-IIIB); about 13 (10-15) furrows on the outer surface at the base. Pecten mandibularis with about 10 (9-12) taeniae; Mdt-Mat 25-28 µm; MTR 0.3-0.39.
Cytology: Four polytene chromosomes with the pseudothummi-cytocomplex combination BF, CD, AE, G.
Nucleolus near centromere in arm F, although a nucleolus is developed subterminal in arm G of some Indian specimens, while in others it may just appear as a puff. Arm G partly unpaired at one end, with prominent BR near middle of the arm and another about one third from the other end and somtimes with a subterminal nucleolus.
Polymorphism in arms A, B, C, F and G: ArmC1 occurs in Varanasi while ArmC2 has been found at Jammu.
ArmA1: 1 - 2c, 4 - 9, 2d - 3, 12- 10, 13 - 19 as circumdatus A2, holomelas, ramosus
ArmA2: 1 - 2c, 10 - 12, 3, 14c - 13, 4 - 9, 2ed, 14d - 19 (India)
ArmB1: Puff distal of centre of arm with dark bands distal; nucleolus near centromere.
ArmC1: Large puff about the middle of the arm.
ArmC2: Inversion of about two thirds of the arm as ramosus
ArmD1:
ArmE1: 1 - 2c, 8b - 6, 3 - 2d, 8 - 9, 4 - 5, 12a - 10, 12b - 13 i.e. In5-9 and In9-3 from dorsalis (as ramosus)
ArmF1: possibly 1 - 10, 15 - 11, 16 - 23 as halophilus
ArmG1: prominent BR near middle of arm, another about 1/3 from paired end.