C. species Bakers Beach

Fourth instar larva: a medium sized bathophilus-type.  No measurements of body or ventral tubule lengths.  Gula and frontoclypeus pale in the single known specimen.  Ventral head length (VHL) 263 micron, width of mentum about 0.7 times the VHL.
Mentum (Fig. a) very similar to that of other members of the C. oppositus group.  The teeth are rounded but this may be due to wear; the 4th laterals are slightly reduced, almost to the level of the 5th laterals (type I-II); the c2 teeth are relatively well separated (type IIA), but the division between the c2 and c1 teeth does not reach down to the level of the outer sides of the c2 teeth.
Ventromental plates (Fig. c) separated by about a third of the mentum width; about 3.2 times longer than deep, with about 28-30 striae; VMR 0.30-0.35.
Pecten epipharyngis difficult to see, but appears to have about 9 teeth, possibly type C.  Premandible with the usual two teeth, inner tooth about twice the width of the outer (about 1.8).
Antenna (Fig. b) with relatively long, narrow basal segment, about 4.1 times as long as wide, and about 0.46 of VHL; RO about a third up from the base; AR about 2.04, relative lengths of segments (micron): 120 : 30 : 8 : 14 : 7.
Distance between the antennal bases may be slightly greater than that between the S4 setae, which are separated by 0.9 of the frontoclypeus width at that point.
Mandible (Fig. d) with third inner tooth pigmented and almost fully separated (II-IIIB); 12 taeniae in the pecten mandibularis and 16 furrows on the outer surface near the base of the mandible.

Cytology:  Three polytene chromosomes with the modified pseudothummi-cytocomplex combination: AEG, BF, CD.  Arm G unpaired with a Balbiani ring (BR) near the fusion with arm E and a second near the middle of the arm.
A nucleolus is developed at about group 19 in arm F.
BB A1:    1a-e, 2d - 3e, 1f - 2c, 10a - 11e, 8 - 9, 15 - 13, 7 - 4, 12a-c, 3i-f, 16 - 19                        as oppositus A5
BB B1:    Large puff (group 7) developed towards the distal end of the arm, with the dark bands (group 8) on the distal side.     as oppositus B2
BB C1:    Banding pattern not clear.
BB D1:    Banding pattern not clear
BB E1:    1 - 2d, 7g - 10b, 3e - 2e, 7f - 3f, 10c - 13                                                                              as oppositus E2
BB F1:    1 - 2a, 10 - 2b, 11 - 23                                                                                                                as oppositus F1
BB G1:    fused to the end of arm E, with two obvious BRs.

This species is known from only a single specimen from Bakers Beach, Tasmania (41.1°:S; 146.6°E), AT.57.1 5F, (Jon Martin & B.T.O. Lee) 9.xii.1979.  The larval morphology and the banding patterns of the polytene chromosomes indicate a relationship to C. oppositus.  The separation of the ventromental plates (0.7 of mentum width) appears to be greater than that of the other members of the C. oppositus-group (0.6 or less).  The larval head capsule and the polytene chromosomes are mounted on separate slides.

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Modified: 16 April 2019
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