Nearest neighbor BIN: BOLD:AAJ3011 an unnamed species (Species 5t) from Canada and Greenland
(based on European specimens in Wülker 1991)(NB: Langton & Visser note that there are differences between the male (sensu Wülker) and those shown on Goeghebuer's original figures)
Pupa: (based on Palearctic specimens from Langton & Visser, 2003):
Exuvial length 9.7 mm (7.2-11.8) mm. Color golden brown to dark brown, difference in color between thorax and abdomen not marked, outer margin of anal lobes colored except at bases.
Cephalic tubercles 108x110 - 200x185 µm; frontal setae30-82 µm long, basal ring of thoracic horn 176 x 77 (120x60-200x80) µm; HR 2.28 (2.13-2.50).
Hook row of segment II entire, occupying 0.47-0.53 of the segment width, with about 68-99 hooks. Armament of tergits not extensive, that of tergites II-VI in form of an undivided, usually extensive patch of strong points, the points in each segment increasing in size posteriorly, the point patches increasing in size from II-V; points of VIII mainly isolated and irregularly arranged. Seta D4 of TVI closer to midline than seta D3. Lateral spur of segment VIII with 1-4 stout apiculate teeth. Anal lobes weakly rounded with 110 (66-140) fringe taeniae, and 1(2) dorsal taenia.
Fourth instar larva of the plumosus-type but smaller than C. tenuistylus.
Gular region darkened only at posterior; frontoclypeus pale; ring organ about 0.3 of the distance from the base of antennal segment 1.
Cytology: 4 polytene chromosomes with the thummi arm combination AB, CD, EF, G. Centromeres heterochromatic.
Arm G generally unpaired with a nucleolus near the heterochromatic centromere, the other end forming a fan-like structure terminated by dark bands, 2 Balbiani rings near the nucleolus. A nucleolus is also present in arm C.
Polymorphism known in arms B (Holarctic), D, F & possibly E (Nearctic).
lonA1: 1 - 2c, 10 - 12, 3 - 2d, 9 - 4, 13 - 19 i.e as holomelas, tenuistylus, etc.
lonC1: 1-6b, 11c-8, 15-11d, 6gh, 17a-16, 7d - a, 6f - c, 17b-22 ie. as aberratus, tenuistylus, etc.
lonD1: 1 - 3, 11 - 18d, 7 - 4, 10 - 8, 18g - 24 ie. as tardus, cucini, pilicornis.
lonE1: 1 - 3e, 5 - 10b, 4 - 3f, 10c - 13 i.e. as tenuistylus, etc.
lonF1: 1 - 23 i.e. Standard, as piger, sp. 3b, etc.
lonG1: Heterochromatic centromere, with nearby nucleolus and 2 BRs.
Found: Newfoundland & Labrador - Torngat Mountains National Park (58.4505°N, 62.7982°W) (BOLD)
Northwest Territories - Nailoicho, Nahanni National Park (61.606°N, 125.758°W) (BOLD). Yukon Territory - Ivvavik National Park (69.162°N, 140.155°W); Kluane National Park (60.748°N, 137.513°W) (both BOLD).
Alaska - No name Lake, Haul Road, South Slope.
Morphology and cytology described by Wülker (1991a).