Species 3x C. sp. Le1. of Kiknadze et al., (1996)

Close to C. riihimakiensis - Kiknadze et al. (2016)

Larva possibly a salinarius-type in North America, but Kiknadze et al. (1996b) describe the Russian larvae as bathophilus-type.  Dark gular region and frontoclypeus.
Mentum with rounded teeth, centre tooth relatively broad and c2 teeth well separated (type IIA), fourth laterals only slightly reduced (type I-II).

Cytology:  4 relatively short polytene chromosomes with the thummi arm combination AB, CD, EF, G.  Centromeres recognizably heterochromatic.
Arm G with nucleolus near the centromere, followed by a constriction and a large median Balbiani ring; superficially similar to that of C. cucini.
Nucleoli in the long arms in both C and D; homologues often partially unpaired.  All sequences, other than LE1A2 and LE1F2, have a Holarctic distribution.

LE1A1:    1-2c, 10-12, 3-2d, 9-4, 13-19            ie. as holomelas
LE1A2:    1-2c, 10, 8d-9, 2d-3, 12-11, 8c-4, 13-19
LE1B1:
LE1C1:    1-6b, 11c-8, 15-12, 11h-d, 6gh, 17a-16, 7d-a, 6f-c, 17(a)b-22        i.e. as pilicornis, etc.
LE1D1:    1-3, 11-17, 8-4, 10-9, 18-24
LE1E1:    1-3e, 5-10b, 4-3f, 10c-13                ie. as aberratus, cucini, etc.
LE1F1:    1-10, 17-11, 18-23                        ie. as aberratus, riihimakiensisF1, etc.
LE1F2:    1-8c, 12-17, 10-8d, 11, 18-23            ie. as riihimakiensisF2
Le1G1:    Subterminal N, median BR.

Found: Nunavut (was North West Territories) - Permanent Pond #78, Hazen, Ellesmere Island (81.82; -71.17) (Oliver & Corbet, 1966).
          Alaska - Point Barrow, Pond B.R.
          Russia - Lena delta.

Karyotype described by Kiknadze et al. (1996b) and redescribed by Wülker & Martin (2000), both as Sp. Le1.

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Modified: 15 March 2021
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