Species k.

Adult not certain.

Pupa:  Caudolateral spur of segment VIII (below) with a single long spine.

Fourth instar larva a small to medium plumosus-type (Fem. abt 11.4 mm (1)).  Ventral tubules moderately long.  Anal tubules about 253-304µm long, ventral pair perhaps longer, and at least twice as long as wide.  Gular region slightly darkened on posterior third, frontoclypeus pale to very slightly dark.  Distance between antennal bases greater than that between S4 setae, which are separated by about 80% of FC width.
Mentum (Fig. c) with pointed teeth; centre tooth high and almost parallel sided, side teeth moderately developed (type IB), 4th laterals hardly reduced (type I).
Ventromentum (Fig. e) about 3.25-3.5 times longer than the depth, with about 41-48 striae reaching about half way to margin; VMR about 0.30-0.33.  Pecten epipharyngis (Fig. a) with about 9-10 moderately long and fairly broad teeth.
Premandible (Fig. b) with relatively broad, about equal length teeth, inner tooth about 1.9-2.3 times wider than outer tooth.
Antenna (Fig. d) with basal segment about 3.7-4 times longer than wide; RO about 0.4-0.5 up from base of segment; AR abt 1.70-1.85; segment proportions (microns): 131 : 40 : 11 : 15 : 8.
Mandible (Fig. f) with 3rd inner tooth only partly separated and pale (type I-IIA), about 12-16 furrows on outer surface near the base, pecten mandibularis with about 13-16 setae.

Cytology:  4 polytene chromosomes with thummi arm combination AB, CD, EF, G.  Centromeres only slightly heterochromatic.
Arm G generally unpaired, with a large terminal nucleolus and at least one Balbiani ring near the other end; rather similar to arm G of C. plumosus.  No nucleolus in any other arm.
Arm E generally unpaired.

Arm A1:    Olive (groups 4-6) about middle of arm
Arm B1:    slight distal puff (group 7?).
Arm B2:    Inversion in proximal half of arm. Arm C1:    typical groups 3-4 about 1/3 from centromere.
Arm D1:
Arm E:    Lack of pairing may be a small inversion near distal end.
Arm F1:
Arm G1:    Large terminal nucleolus, one or 2 BR near other end.

Found: Ontario - Costello Creek, Algonquin Provincial Park (45.58°N; 78.32°W).
            New Hampshire - Lancaster, Coos Co. (44.49°N; 71.57°W).

            Creeks and pools.

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Modified: 31 January 2019
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