Species x. C. hyperboreus Staeger, 1845

Molecular sequence from Canada is under the name C. species TE13.

In Bold Bin: BOLD:AAC0596

Adult

Male: (from Townes, 1945) and photo in BOLD):


C. hyperboreus male from BOLD (left) and hypopygium from Townes 1945 (right)

Wing length 4.2-5.2 mm; fore LR 1.25-1.27; antennal ratio 5.0.
Body moderately stout.
Frontal tubercles rather small, clypeus small.
Middle portion of pronotum slightly broadened; mesoscutum without a tubercle.
Thorax, scutellum, etc., blackish, abdominal segments blackish at base, with a brownish band across the distal ¼ to 1/3; bases of anterior and midlegs brownish, rest of legs blackish.
Fore tarsus with a long dense beard, BR about 4.
Townes notes that the specimen described has paler legs than the types.
Leg proportions (micron)

 
Fe
Ti
Ta1
Ta2
Ta3
Ta4
Ta5
LR
F/T
BR
PI
1200
1440
1720
710
550
475
400
1.20
0.83
abt 4
PII
1000
1640
880
720
360
280
200
0.54
0.63
 
PIII
2000
2200
-
-
-
-
-
-
0.91
 

Townes shows about 11 setae in a number of pale patches on TIX. Anal point narrow, extending beyond midpoint of gonostyle; Superior volsella rather unusual, not easily fitted into the Strenzke groups, closest to E(i) but more curved and of similar width up to its distal end. Inferior volsella not reaching to end of anal point, but to about midpoint of gonostylus.

Female:

From a photograph in the BOLD database (above), the following characters can be adduced:
Wing length about 3.1 mm.
Thorax, scutellum, etc, blackish, abdominal segmets blackish at base, with a brownish band across the distal ¼ to 1/3; bases of anterior and midlegs brownish, rest of legs blackish.

 
Fe
Ti
Ta1
Ta2
Ta3
Ta4
Ta5
LR
F/T
Ta5/Ti
PI
885
914
-
-
-
-
-
-
0.97
0.21
PII
940
-
-
-
-
-
-
-
-
 
PIII
1157
1343
1050
514
385
214
157
0.78
0.86
 

Pupa: no information available.

Larva: a medium sized (male 12.2-14.5 mm) salinarius-type.  Anal tubules relatively long (dorsal 375x140, ventral 401x185 µm), dorsal about 3 times longer than wide, the ventral pair about 2.2 times longer.
Gular region, frontoclypeus and sometimes other parts of the head, darkened. Oesophageal opening about 61-78 µm x 18-25 ݙm (l/w 3.1-3.4).

Mentum of all specimens badly worn, but c2 teeth are well seperated (type IIA) and fourth laterals reduced (type II).
Ventromentum abt 2.05 wide and abt 3.3 times wider than deep; sl. shorter to same width as mentum; abt 42-44 striae; VMR about 9.24.
Pecten epipharyngis with 8-15 teeth, sometimes irregular and broad.
Premandible with broad teeth about equal in length; inner tooth about 3-4 times wider than outer tooth.
Antenna with basal segment about 3.1-3.3 times longer than wide, RO about 0.40-0.55 up from base; AR about 1.87-1.94; A2 about 0.29-0.31 of A1; segment proportions (micron) 126 : 37 : 11 : 14 : 7.5.
Distance between antennal bases about the same as the distance between the S4 setae, which are separated by about 0.78 of the FC width.
Mandible abt 245-266 µm long, with 3rd inner tooth usually well developed (type IIB), but may be broken, and with about 14-16 furrows on the outer surface near the base, PecM abt 13-14; Mdt-Mat abt 25 µm, MTR 0.28.

Cytology:  4 short polytene chromosomes with the thummi arm combination AB, CD, EF, G, but Keyl pattern hard to recognise.
Centromeres heavily heterochromatic, often forming a chromocentre.
Arm G short, often unpaired, with a nucleolus near the centromere and possibly a Balbiani ring near the other end.
Polymorphic in arm A.  Polymorphism also occurs in arm F, but only F2 is present at Ellesmere.

hypA1:    1 - 2c, 10 - 12, 3 - 2d, 9 - 4, 13 - 19    ie. as holomelas A1
hypA2:    1 - 2c, 10 - 12, 7 - 9, 2d - 3, 6 - 4, 13 - 19    (Greenland)
hypA3:    1 - 2c, 3 - 2d, 9 - 7, 12 - 10, 6 - 4, 13 - 19    (Ellesmere)
hypB1:    as riihimakiensis
hypC1:    1-2c, 3-2d, 9-7, 12-10, 6-4, 13-19    (Wülker)    or
                  1-6b, 11c-8, 15-11d, 6gh, 17a, 16h-a, 7d-a, 6f-c, 17b-22    (Kiknadze)
hypD1:    1 - 3, 11 - 18d, 7 - 4, 10 - 7e, 18g - 24    ie. as longistylus, tenuistylus, etc.
hypE1:    1 - 3e, 7 - 5, 8 - 10b, 4 - 3f, 10c - 13
hypF1:    1 - 6, 12 - 7, 13 - 23    (Greenland)
hypF2:    1 - 2, 14 - 13, 7 - 12, 6 - 3, 15 - 23

Found: Alberta - NE Jasper Lake, Jasper National Park (as sp. TE13)
            Nunavut (formerly North West Territories) - Hazen Camp Pond, Skeleton Lake, and its inlet marsh, Hazen, Ellesmere Island
            (Oliver & Corbet, 1966).
            Greenland - Egedesminde (now Aasiaat)(68.71°N, 52.87°W) (Townes 1945); Lake 517, Stoe Kvaneso, Western Greenland
           (Type locality); Ilimaussaq intrusion, South Greenland (60-61°N, 45-46°W) (Wülker & Butler 1983); Nedre Midsommer Sö, Peary Land
            (82.63°N, 32.50°W); Zackenberg Research Station, Northeast Greenland (GenBank).

            Arctic lakes and pools.

Chromosomes pictured and briefly described by Wülker & Butler (1983), redescribed by Wülker & Martin (2000).

Molecular sequences:
mtCOI:  A number of sequences are available in GenBank or BOLD, under the names C. hyperboreus or C. sp. TE13 (from Arctic Canada).

See also C. rempeli and C. sp. u

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Modified: 24 August 2018
Access: Unrestricted
Copyright © 2000-2018, Jon Martin.