Pupa: Only the caudolateral spur of segment VIII has been recorded, with 1-3 spines at the end of the spur only.
Fourth instar larva a medium sized plumosus-type. No metrics recorded for tubuli.
Gular region darkened over posterior half, frontoclypeus probably darkened.
Mentum (Fig. c) with pointed teeth; c1 tooth relatively broad with diverging sides, c2 teeth, although well seperated, virtually continue line of centre tooth (type 1B or III). Fourth laterals not reduced (type I).
Ventromental plates (Fig. d) separated by about 0.36 of mentum width, with about 40-45 striae, VMR about 0.25-0.33. Pecten epipharyngis (Fig. a) with about 10-11 sharp but relatively broad teeth (type B).
Premandible with outer tooth longer, inner tooth about three times broader than outer.Antenna (Fig. b) with basal segment about 3.5-3.9 times longer than wide, RO nearly halfway up from base of segment; AR abt 2.1-2.33; relative lengths of segments (micron) 140 : 30 : 8 : 13 : 8.
Mandible (Fig. e) with third inner tooth well developed but only partly darkened (Type IIIB), about 16 furrows on outer surface near the base; 12-13 setae in Pecten mandibularis.
Cytology: 4 polytene chromosomes with the thummi arm combination AB, CD, EF, G.
Centromeres heterochromatic, showing tendency to stick together to form a centromere.
Large Balbiani ring (BR) near centre of closely paired arm G, with constriction between BR and centromere; another BR distal in the arm.
Nucleolus in arm B, just distal to the 4 characteristic bands; 1 or 2 nucleoli near the middle of arm D.
No polymorphism in small sample examined.
Arm A1: 1a-e, 2d-3, 12-10, 14 - 13, 4 - 9, 1f- 2c, 15-19
Arm B1: Nucleolus about 1/3 from centromere
Arm C1: possibly 1 - 6b, 11c - 8, 15 - 11d, 6gh, 17a - 16, 7d - 6c, 17b - 22 i.e. as sp. Apple Valley (sp. 3b)
Arm D1: 1 or 2 nucleoli near middle of arm
Arm E1: 1 - 3e, 5 - 10b, 4 - 3f, 10 - 13 i.e. as cingulatus, tardus and sp. 3b.
Arm F1: 1 - 23 i.e. Standard, as piger
Found: California - Lake Davis, Plumas Co. (abt 39.70°N, 120.50°W).
Some information on arm F given in Fig. 3 of Martin (1979).